Two nights ago, I attended a meeting of the Los Angeles Bead Society. Jamey Allen was the speaker and the topic was Beads From Borneo. Many of the types of beads that are in your photo were discussed. I own numerous strands from Borneo, including some of the types in your photo. It is my opinion that most of the beads -most, but not all - are Venetian. You raise an interesting point, that the beads are smaller than the trade beads that we know from West Africa. Smaller Venetian beads are also found in Peru, Bolivia and Mexico.